
In terrestrial vertebrates, digitigrade (/ˈdɪdʒɪtɪˌɡreɪd/) locomotion is walking or running on the toes (from the Latin digitus, 'finger', and gradior, 'walk'). A digitigrade animal is one that stands or walks with its toes (metatarsals) touching the ground, and the rest of its foot lifted. Digitigrades include walking birds (what many assume to be bird knees are actually ankles), cats, dogs, and many other mammals, but not plantigrades or unguligrades. Digitigrades generally move more quickly and quietly than other animals.